1. Department of Ecology, School of Life SciencesSun Yat‐sen University Guangzhou Guangdong China
2. Institute of Eastern–Himalaya Biodiversity ResearchDali University Dali Yunnan China
3. Department of Genetics, Evolution and Environment School of Life and Medical SciencesUniversity College London London United Kingdom
4. Institute of ZoologyZoological Society of London London United Kingdom