1. U.S. Geological Survey, Forest and Rangeland Ecosystem Science Center 600 East Park Ave Port Angeles WA 98362 USA
2. U.S. Geological Survey, National Wildlife Health Center Madison WI 53711 USA
3. U.S. Geological Survey, Northern Rocky Mountain Science Center, Missoula, MT 59801, USA and Wildlife Biology Program, W. A. Franke College of Forestry and Conservation, University of Montana Missoula MT 59801 USA