1. Department of Biology and Biotechnologies “Charles Darwin” University of Rome La Sapienza Rome Italy
2. CEFE, CNRS University of Montpellier, University Paul Valéry Montpellier 3, EPHE, IRD Montpellier France
3. Italian Institute for Environmental Protection and Research (ISPRA), Unit for Conservation Genetics (BIO–CGE), Ozzano dell'Emilia Italy
4. Wolf Apennine Center, Appennino Tosco‐Emiliano National Park, Ligonchio Italy