1. Aquatic Systems Research Group, Department of Geography and Environmental Sciences University of Venda Thohoyandou South Africa
2. GEOMAR Helmholtz‐Zentrum für Ozeanforschung Kiel Kiel Germany
3. South African Institute for Aquatic Biodiversity Makhanda South Africa
4. Department of Zoology and Entomology Rhodes University Makhanda South Africa
5. Aquatic Systems Research Group, School of Biology and Environmental Sciences University of Mpumalanga Nelspruit South Africa