1. The Royal Veterinary College; Equine Referral Hospital; Hawkshead Lane Hatfield AL9 7TA
2. Department of Musculoskeletal Biology; Institute of Ageing and Chronic Disease; The University of Liverpool; William Henry Duncan Building West Derby Street Liverpool Merseyside L7 8TX
3. Institute of Veterinary Sciences; The University of Liverpool, Leahurst; Chester High Road Neston Cheshire CH64 7TE