1. Department of Genetics and Genome Biology The University of Leicester, Leicester LE1 7RH United Kingdom
2. Laboratory of Socioecology and Social Evolution Department of Biology, KU Leuven, 3000 Leuven Belgium
3. Biobest Belgium N.V., 2260 Westerlo Belgium
4. The Lancaster Environmental Centre University of Lancaster, Lancaster LA1 4YW United Kingdom