1. School of Anatomical Sciences Faculty of Health Sciences, University of the Witwatersrand 7 York Road, Parktown, 2193 Johannesburg Republic of South Africa
2. Division of Clinical Anatomy and Biological Anthropology, Department of Human Biology University of Cape Town Cape Town South Africa
3. Centre for Zoo and Wild Animal Health, Copenhagen Zoo Frederiksberg Denmark
4. Borås Zoo Borås Sweden
5. Department of Anthropology and Center for the Advanced Study of Human Paleobiology The George Washington University Washington DC USA