1. Department of Chemical Science and Engineering Tokyo Institute of Technology 2-12-1-H101 Ookayama, Meguro Tokyo 152-8552 Japan
2. Division of Molecular Immunology, Medical Institute of Bioregulation Kyushu University 3-1-1 Maidashi, Higashiku Fukuoka 812-8582 Japan
3. Current address: Department of Molecular Immunology, Research Institute for Microbial Diseases, and Laboratory for Molecular Immunology Osaka University 3-1 Yamadaoka, Suita 565-0871, Osaka University Japan
4. Current address: WPI Immunology Frontier Research Center (iFReC) Osaka University 3-1 Yamadaoka Suita 565–0871 Japan
5. Department of Ophthalmology, Graduate School of Medical Sciences Kyushu University 3-1-1 Maidashi, Higashiku Fukuoka 812-8582 Japan