1. Department of Ecosystem Studies, School of Environmental Science The University of Shiga Prefecture Hikone Shiga Japan
2. Creative Science Unit (Geoscience), Faculty of Science Shizuoka University Suruga‐ku Shizuoka Japan
3. Lake Biwa Environmental Research Institute Otsu Shiga Japan
4. Shiga Prefectural Fisheries Experiment Station Hikone Shiga Japan