1. Netherlands Center for Biodiversity Naturalis (section NHN); Leiden University; P.O. Box 9514 2300, RA Leiden Netherlands
2. Utrecht University; Laboratory of Palaeobotany & Palynology; Budapestlaan 4 3584 CD Utrecht Netherlands
3. Wageningen University; Biosystematics group; Droevendaalsesteeg 1 6708 PB Wageningen Netherlands