1. Heilongjiang Provincial Key Laboratory of Ecological Restoration and Resource Utilization for Cold Region School of Life Sciences Heilongjiang University Harbin China
2. Institution of Nature and Ecology Heilongjiang Academy of Sciences Harbin China
3. Swiss Federal Research Institute WSL Birmensdorf Switzerland
4. Heilongjiang Academy of Forestry Harbin China
5. Key Laboratory of Geographical Processes and Ecological Security in Changbai Mountains Ministry of Education School of Geographical Sciences Northeast Normal University Changchun China
6. CAS Key Laboratory of Forest Ecology and Management Institute of Applied Ecology Erguna Forest‐Steppe Ecotone Research Station Chinese Academy of Sciences Shenyang China