1. Microbiology and Metabolic Engineering Key Laboratory of Sichuan Province; College of Life Science, Sichuan University; Chengdu Sichuan 610065 People's Republic of China
2. Department of Microbiology and Immunology; Institute of Medical and Pharmaceutical Sciences, Zhengzhou University; Zhengzhou 450052 People's Republic of China
3. National Engineering Research Center for Biomaterials; Sichuan University; Chengdu 610065 People's Republic of China