1. Department of Botany and Plant Sciences University of California 2150 Batchelor Hall Riverside CA92521USA
2. Evolution, Ecology and Organismal Biology Graduate Program Department of Evolution, Ecology and Organismal Biology University of California Riverside CA92521USA
3. Centro de Investigaciones Biológicas del Noroeste Av. Instituto Politécnico Nacional 195, Col. Playa Palo de Santa Rita Sur La Paz B.C.S23096México
4. Smithsonian Tropical Research Institute Apartado 0843‐03092. Balboa, Ancon Panama Republic of Panama