1. Department of Biological & Environmental Sciences Georgia College and State University Milledgeville Georgia31062‐0001USA
2. Department of Biology University of Rwanda Huye Rwanda
3. The Dian Fossey Gorilla Fund International Karisoke Research Center Ruhengeri Rwanda
4. University of Rwanda Kigali Rwanda
5. School of Life Sciences Arizona State University PO Box 874501 Tempe Arizona85287‐4501USA