1. Kunming Institute of Botany Chinese Academy of Sciences Kunming Yunnan 650204 China
2. Department of Botany Smithsonian Institution Washington D.C. 20013 USA
3. Florida Museum of Natural History University of Florida Gainesville Florida 32611 USA
4. Biodiversity Institute University of Florida Gainesville Florida 32611 USA
5. Department of Biology University of Florida Gainesville Florida 32611 USA
6. Department of Ecology and Evolutionary Biology University of Michigan Ann Arbor Michigan 48109 USA