1. ARC Centre of Excellence in Vision Science; Australian National University; Canberra ACT 0200 Australia
2. Research School of Biology; College of Medicine; Biology and Environment; Australian National University; Canberra ACT 0200 Australia
3. Centre for Integrative Ecology; School of Life and Environmental Sciences; Deakin University; Geelong VIC 3217 Australia
4. Medical School and the John Curtin School of Medical Research; College of Medicine; Biology and Environment; Australian National University; Canberra ACT 0200 Australia
5. School of Animal Biology and the UWA Oceans Institute, University of Western Australia; Crawley WA 6009 Australia