1. School of Life Sciences, Key Laboratory of Conservation and Application in Biodiversity of South China Guangzhou University, Guangzhou Higher Education Mega Center Guangzhou PR China
2. School of Earth Sciences and Engineering, Guangdong Provincial Key Lab of Geological Processes and Mineral Resources Sun Yat‐sen University Guangzhou PR China
3. College of Life Sciences and Academy for Multidisciplinary Studies, Capital Normal University Beijing PR China