1. Cancer Research UK Edinburgh CentreMedical Research Council Institute of Genetics & Molecular Medicine, Western General Hospital, The University of Edinburgh Edinburgh United Kingdom
2. Colon Cancer Genetics Group, Medical Research Council Human Genetics UnitMedical Research Council Institute of Genetics & Molecular Medicine, Western General Hospital, The University of Edinburgh Edinburgh United Kingdom
3. Centre for Global Health ResearchUsher Institute of Population Health Sciences and Informatics, The University of Edinburgh Edinburgh United Kingdom