1. Univ Brest, CNRS, IRD, Ifremer, LEMAR F‐29280, Plouzane France
2. Sorbonne University, CNRS, UMR7144, Ecology of Marine Plankton Team, Station Biologique de Roscoff Roscoff France
3. National Institute of Water and Atmospheric Research Wellington New Zealand
4. Centro Oceanográfico de Gijón/Xixón Instituto Español de Oceanografía (IEO) Gijón/Xixón Asturias Spain
5. National Institute of Water and Atmospheric Research Hamilton New Zealand