1. Finnish Environment Institute Helsinki Finland
2. Department of Environmental Sciences University of Helsinki Finland
3. Geological Survey of Finland Kuopio Finland
4. Department of Environmental Sciences University of Eastern Finland Kuopio Finland
5. Department of Biology University of Joensuu Joensuu Finland
6. Department of Forest Sciences University of Helsinki Finland
7. Environmental Studies Program Mount Holyoke College South Hadley, Massachusetts USA
8. Finnish Forest Research Institute Joensuu Finland