1. Faculty of Science, Department of Biology; Yamagata University; Kojirakawa Yamagata Japan
2. Graduate School of Engineering; Utsunomiya University; Utsunomiya Tochigi Japan
3. Department of Biology; University of Oulu; Oulu Finland
4. Research Institute of Luminous Organisms; Hachijojima Tokyo Japan
5. Institute of Arts and Sciences; Yamagata University; Yamagata Japan