1. Makino Herbarium Tokyo Metropolitan University Minamiosawa 1‐1 Hachioji Tokyo 192‐0397 Japan
2. Department of Biology, Faculty of Science Chiba University Yayoi‐cho Inage Chiba 263‐8522 Japan
3. RIKEN Interdisciplinary Theoretical and Mathematical Sciences (iTHEMS) Wako Saitama 351‐0198 Japan
4. The Kochi Prefectural Makino Botanical Garden 4200‐6 Godaisan Kochi 781‐8125 Japan
5. Department of Botany National Museum of Nature and Science 4‐1‐1 Amakubo Tsukuba Ibaraki 305‐0005 Japan
6. Department of Biology, Graduate School of Science Chiba University Yayoi‐cho Inage Chiba 263‐8522 Japan