1. Université de Rennes 1; Ethologie animale et humaine; UMR 6552 - CNRS Paimpont France
2. School of Biology; Indian Institute of Science, Education and Research, Thiruvananthapuram; Thiruvananthapuram India
3. Department of Zoology; Institute of Ecology and Earth Sciences; University of Tartu; Tartu Estonia
4. Institut universitaire de France; Paris France