1. School of Environment and Sustainability Western Colorado University Gunnison Colorado 81231 USA
2. Department of Biology Utah State University Logan Utah 84322 USA
3. Department of Forest Ecosystems and Society Oregon State University Corvallis Oregon 97331 USA
4. Haire Laboratory for Landscape Ecology Belfast Maine 04915 USA
5. Aldo Leopold Wilderness Research Institute Rocky Mountain Research Station USDA Forest Service Missoula Montana 59801 USA
6. Northern Forestry Centre Canadian Forest Service Natural Resources Canada Edmonton Alberta Canada