1. Guangdong Entomological Institute South China Institute of Endangered Animals Guangzhou 510260 China
2. Institute of Genetic Resources College of Life Sciences Nanjing Normal University Nanjing 210046 China
3. State Key Laboratory of Biocontrol and School of Life Sciences SunYat‐sen University Guangzhou 510275 China
4. State Key Laboratory of Genetic Resources and Evolution Kunming Institute of Zoology Chinese Academy of Sciences Kunming 650223 China
5. Ministry of Education Key Laboratory for Biodiversity Sciences and Ecological Engineering College of Life Sciences Beijing Normal University Beijing 100875 China