1. Montana Cooperative Fishery Research Unit Department of Ecology Montana State University Post Office Box 173460 Bozeman Montana 59717 USA
2. U.S. Geological Survey Montana Cooperative Fishery Research Unit Department of Ecology Montana State University Post Office Box 173460 Bozeman Montana 59717 USA
3. U.S. National Park Service Yellowstone Center for Resources Native Fish Conservation Program Post Office Box 168, Yellowstone National Park Wyoming 82190 USA