1. Fish Ecology and Conservation Physiology Laboratory Department of Biology Carleton University 1125 Colonel By Drive Ottawa Ontario K1S 5B6 Canada
2. Great Lakes Laboratory for Fisheries and Aquatic Sciences, Fisheries and Oceans Canada Burlington Ontario L7S 1A1 Canada
3. Department of Environmental Conservation University of Massachusetts Amherst 160 Holdsworth Way Amherst Massachusetts 01003 USA