1. Fish Ecology and Conservation Physiology Laboratory Department of Biology and Institute of Environmental and Interdisciplinary Science Carleton University Ottawa Ontario K1S 5B6 Canada
2. The Kenauk Institute Kenauk Nature 1000 Chemin Kenauk Montebello Quebec J0V 1L0 Canada
3. Fisheries and Oceans Canada Great Lakes Laboratory for Fisheries and Aquatic Sciences Burlington Ontario L7S 1A1 Canada
4. Department of Environmental Conservation University of Massachusetts Amherst Amherst Massachusetts 01003 USA