1. Department of Biology Trent University Peterborough Ontario K9L 1Z8 Canada
2. Environmental and Life Sciences Graduate Program Trent University Peterborough Ontario K9L 1Z8 Canada
3. Ontario Ministry of Northern Development, Mines, Natural Resources and Forestry, Aquatic Research and Monitoring Section, Codrington Fisheries Research Facility Codrington Ontario K0K 1R0 Canada
4. Ontario Ministry of Northern Development, Mines, Natural Resources and Forestry, Aquatic Research and Monitoring Section Trent University Peterborough Ontario K9J 7B8 Canada