1. Okinawa Institute of Science and Technology Graduate University (OIST) 1919-1 Tancha, Onna-son; Kunigami-gun Okinawa Japan
2. National Institute of Oceanography and Fisheries; Cairo Egypt
3. Division of Biological and Environmental Sciences and Engineering; Red Sea Research Center; King Abdullah University of Science and Technology; Thuwal Saudi Arabia
4. Department of Ecology and Evolutionary Biology; University of California Santa Cruz; Santa Cruz CA USA