1. Department of Zoology, Entomology and Fisheries Sciences, College of Natural Sciences Makerere University Kampala Uganda
2. National Forestry Resources Research Institute Mukono Uganda
3. Charles Perkins Centre and School of Life and Environmental Sciences The University of Sydney Sydney New South Wales Australia
4. Department of Forestry, Biodiversity and Tourism, College of Agricultural and Environmental Sciences Makerere University Kampala Uganda
5. Department of Human Evolutionary Biology Harvard University Cambridge Massachusetts USA
6. Department of Anthropology Hunter College of the City University of New York New York City New York USA