1. Evolutionary Morphology Section, Department of Evolution and Phylogeny; Primate Research Institute, Kyoto University; Japan
2. Department of Zoology; Mawlamyine University; Mawlamyine Myanmar
3. Department of Biology, Faculty of Science; Chulalongkorn University; Bangkok Thailand
4. National Primate Research Center of Thailand; Chulalongkorn University; Saraburi Thailand