1. NIHR Newcastle Biomedical Research Centre; Newcastle upon Tyne Hospitals NHS Foundation Trust, and Newcastle University; Newcastle upon Tyne UK
2. Arthritis Research UK Centre for Genetics and Genomics; Centre for Musculoskeletal Research; Institute of Inflammation and Repair; University of Manchester; and NIHR Manchester Musculoskeletal Biomedical Research Unit; Central Manchester NHS Foundation Trust; Manchester UK
3. Newcastle University; Newcastle upon Tyne UK