1. Department of Gastrointestinal Surgery College of Life Sciences Zhongnan Hospital of Wuhan University Wuhan University Wuhan 430071 China
2. Department of Immunology Medical Research Institute and Frontier Science Center for Immunology and Metabolism Wuhan University Wuhan 430071 China
3. Wuhan Research Center for Infectious Diseases and Cancer Chinese Academy of Medical Sciences Wuhan 430071 China
4. State Key Laboratory of Virology Hubei Key Laboratory of Cell Homeostasis College of Life Sciences Frontier Science Center for Immunology and Metabolism Wuhan University Wuhan 430072 China
5. The Institute for Advanced Studies Wuhan University Wuhan 430072 China