1. School of Life Science and Technology ShanghaiTech University Shanghai 201210 China
2. CAS Center for Excellence in Molecular Cell Science, Shanghai Institute of Biochemistry and Cell Biology Chinese Academy of Sciences Shanghai 200031 China
3. University of Chinese Academy of Sciences Beijing 100049 China
4. Interdisciplinary Research Center on Biology and Chemistry, Shanghai Institute of Organic Chemistry Chinese Academy of Sciences Shanghai 201210 China
5. College of Life Sciences Northwest A&F University Yangling Shaanxi 712100 China
6. Department of Physics, Laboratory of Atomic and Solid State Physics Cornell University Ithaca NY 14853 USA
7. Howard Hughes Medical Institute Cornell University Ithaca NY 14853 USA