1. Department of Life Science College of Natural Sciences Hanyang University Research institute for Natural Sciences Hanyang University Seoul 04763 Republic of Korea
2. Hubrecht Institute for Developmental Biology and Stem Cell Research‐KNAW, University Medical Centre Utrecht Utrecht 3584 CT Netherland
3. Department of Neurology College of Medicine Seoul National University Seoul National University Hospital Seoul 03080 Republic of Korea
4. Department of Life Science College of Natural Sciences Hanyang University Research institute for Natural Sciences Research Institute for Convergence of Basic Sciences Hanyang University Seoul 04763 Republic of Korea