1. Institute for Immunology Tsinghua University Beijing 100084 China
2. Department of Basic Medical Sciences School of Medicine Tsinghua University Beijing 100084 China
3. Beijing Key Lab for Immunological Research on Chronic Diseases Tsinghua University Beijing 100084 China
4. CAS Key Laboratory of Tissue Microenvironment and Tumor Shanghai Institute of Nutrition and Health Chinese Academy of Sciences Shanghai 200031 China