1. School of Pharmaceutical Sciences Tsinghua University Beijing 100084 China
2. Beijing Institutes of Life Science Chinese Academy of Sciences University of Chinese Academy of Sciences Beijing 100101 China
3. State Key Laboratory of Molecular Oncology National Cancer Center/National Clinical Research Center for Cancer/Cancer Hospital Chinese Academy of Medical Sciences and Peking Union Medical College Beijing 100021 China
4. Department of Colorectal Surgery National Cancer Center/National Clinical Research Center for Cancer/Cancer Hospital Chinese Academy of Medical Sciences and Peking Union Medical College Beijing 100021 China
5. Department of Pathology National Cancer Center/National Clinical Research Center for Cancer/Cancer Hospital Chinese Academy of Medical Sciences and Peking Union Medical College Beijing 100021 China
6. Key Laboratory of Systems Biology Hangzhou Institute for Advanced Study University of Chinese Academy of Sciences Hangzhou 310024 China
7. Center for Excellence in Animal Evolution and Genetics Chinese Academy of Sciences Kunming 650223 China