1. University of Tunis El Manar; Biology department; Tunish Tunisia
2. Department of Virology; Amiens University Medical Centre; Amiens France
3. Virology Research Unit, EA 4294; Jules Verne University of Picardie; Amiens France
4. Faculty of Sciences of Tunis; department of Biology, Laboratory of Mycology, Pathologies and Biomarkers; Tunis Tunisia
5. School of Medicine; Nazarbayev University; Astana Kazakhstan