1. Graduate School of Life Sciences Tohoku University 2-1-1 Katahira Aoba-ku, Sendai 980-8577 Japan
2. Cell Biology Center Institute of Innovative Research Tokyo Institute of Technology 4259-S2-19, Nagatsuta-cho Midori-ku Yokohama Kanagawa 226-8503 Japan
3. Laboratory for Chemistry and Life Science Institute of Innovative Research Tokyo Institute of Technology 4259-R1-13 Nagatsuta-cho Midori-ku Yokohama Kanagawa 226-8503 Japan
4. Frontier Research Institute for Interdisciplinary Sciences Tohoku University 2-1-1 Katahira Aoba-ku, Sendai 980-8577 Japan