1. Department of Soil and WaterEEAD‐CSIC Avda. Montañana 1005 Zaragoza 50059 Spain
2. Institute for Biodiversity and Ecosystem Dynamics (IBED)University of Amsterdam PO Box 94240 Amsterdam 1090 GE The Netherlands
3. Department of Geography, Environmental Sciences InstituteUniversity of Zaragoza Pedro Cerbuna 12 Zaragoza 50009 Spain