1. Aix Marseille Université, CNRS/INSU, Université de Toulon, IRD, Mediterranean Institute of Oceanography (MIO) UM 110 Marseille France
2. Laboratoire des sciences de l'environnement marin, IUEM Université de Brest‐UMR 6539 CNRS/UBO/IRD/Ifremer Plouzané France
3. Aix Marseille Université, CNRS/INSU, Université de Toulon, IRD, Mediterranean Institute of Oceanography (MIO) UM 110 New Caledonia France
4. Marine Biological Section, Department of Biology University of Copenhagen Helsingør Denmark
5. Department of Biology University of Massachusetts Dartmouth North Dartmouth Massachusetts USA