1. Department of Orthopaedic Surgery; UCLA-Orthopaedic Hospital; Los Angeles CA 90095 United States
2. Department of Molecular, Cell and Developmental Biology; University of California, Los Angeles; Los Angeles CA 90095 United States
3. Mount Desert Island Biological Laboratory; 159 Old Bar Harbor Road Salisbury Cove ME 04672 United States
4. Center for Endocrinology, Diabetes and Metabolism, Institute of Biomedical Research; University of Birmingham; Birmingham B15 2TT UK