1. Institute of Marine Science and TechnologyShandong University Jinan China
2. Research Centre for Indian Ocean EcosystemTianjin University of Science and Technology Tianjin China
3. Tianjin Key Laboratory of Marine Resources and ChemistryTianjin University of Science and Technology Tianjin China
4. State Key Laboratory of Tropic OceanographySouth China Sea Institute of OceanologyChinese Academy of Sciences Guangzhou China