1. Graduate School of Agricultural and Life Sciences University of Tokyo 1‐1‐1 Yayoi, Bunkyo‐ku Tokyo113‐8657Japan
2. National Institute for Environmental Studies Center for Health and Environmental Risk Research 16‐2 Onogawa Tsukuba Ibaraki305‐8506Japan
3. Faculty of Environment and Information Sciences Yokohama National University 79‐7 Tokiwadai, Hodogaya‐ku Yokohama240‐8501Japan
4. Environmental and Community Affairs Department Nature Conservation Division Chiba Biodiversity Center Chiba Prefectural Government 955‐2 Aoba‐cho, Chuo‐ku Chiba City Chiba260‐8682Japan
5. Japan Wildlife Research Center 3‐3‐7 Kotobashi, Sumida‐ku Tokyo130‐8606Japan
6. Faculty of Science Toho University 2‐2‐1 Miyama Funabashi Chiba274‐8510Japan
7. National Institute for Environmental Studies Center for Environmental Biology and Ecosystem Studies 16‐2 Onogawa Tsukuba Ibaraki305‐8506Japan