1. Department of Ecology and Genetics University of Oulu Oulu Finland
2. Natural Resources Institute Finland Oulu Finland
3. Center for Macroecology, Evolution and Climate GLOBE Institute University of Copenhagen Copenhagen Denmark
4. ISEM, Univ Montpellier, CNRS, EPHE, IRD Montpellier France
5. Department of Biological and Environmental Science University of Jyväskylä Jyväskylä Finland
6. Finnish Museum of Natural History Helsinki Finland
7. National Marine Fisheries Service, NOAA Seattle Washington USA