1. Department of Ecology and Evolutionary Biology Brown University Box G‐W, 80 Waterman Street Providence Rhode Island 02912 USA
2. Department of Biological Sciences University of Arkansas Fayetteville Arkansas 72701 USA
3. School of Biology and Ecology University of Maine Orono Maine 04469 USA
4. Department of Plant Sciences University of California, Davis 1 Shields Avenue Davis California 95616 USA
5. Department of Biological Sciences Central Washington University 400 E. University Way Ellensburg Washington 98926 USA
6. CSIRO National Research Collections Australia Australian National Herbarium Canberra ACT 2601 Australia