1. Department of Medicinal Plants and Cultivation, College of Pharmacy Guizhou University of Traditional Chinese Medicine Guiyang 550025 China
2. Laboratory of Subtropical Biodiversity Jiangxi Agricultural University Nanchang 330045 China
3. Key Laboratory of Forest Plant Cultivation, Development and Utilization of Yunnan Province, Yunnan, China; Yunnan Key Laboratory of Conservation and Breeding of Rare and Endangered Forest Plants, State Forestry Administration, Yunnan Academy of Forestry and Grassland Science Kunming 650201 Yunnan China
4. Key Laboratory for Plant Diversity and Biogeography of East Asia Kunming Institute of Botany, Chinese Academy of Sciences Kunming 650201 China
5. CAS Key Laboratory of Mountain Ecological Restoration and Bioresource Utilization; Ecological Restoration and Biodiversity Conservation Key Laboratory of Sichuan Province, Chinese Academy of Sciences Chengdu Institute of Biology Chengdu Sichuan 610041 China
6. Yunnan Academy of Forestry and Grassland Science Kunming 650201 Yunnan China