1. College of Marine Sciences Shanghai Ocean University Shanghai 201306 China
2. College of Marine Sciences Shanghai Ocean University National Engineering Research Center for Oceanic Fisheries Key Laboratory of Sustainable Exploitation of Oceanic Fisheries Resources Ministry of Education, and Key Laboratory of Oceanic Fisheries Exploration and Scientific Observing and Experimental Station of Oceanic Fishery Resources Ministry of Agriculture and Rural Affairs Shanghai 201306 China
3. College of Marine Sciences Shanghai Ocean University National Engineering Research Center for Oceanic Fisheries Key Laboratory of Sustainable Exploitation of Oceanic Fisheries Resources Ministry of Education Key Laboratory of Oceanic Fisheries Exploration Ministry of Agriculture and Rural Affairs Shanghai 201306 China