1. Department of Wildlife, Fisheries and Conservation Biology University of Maine 5755 Nutting Hall Orono Maine 04469 USA
2. National Oceanic and Atmospheric Administration Northeast Fisheries Science Center, Resource Evaluation and Assessment/Protected Species Woods Hole Massachusetts 02543 USA
3. The Nature Conservancy 2101 North Front Street, Building 1, Suite 200 Harrisburg Pennsylvania 17110 USA
4. U.S. Geological Survey Maine Cooperative Fish and Wildlife Research Unit, and Department of Wildlife, Fisheries and Conservation Biology University of Maine 5755 Nutting Hall Orono Maine 04469 USA